Born on July 26, 1973 in London, England, Beckinsale pursued her education at Godolphin & Latymer School, then Oxford University, where she took up acting. In "Royal Deceit/The Prince of Jutland" (1994), which was based on the Danish prince whose life inspired Shakespeare's "Hamlet,” she starred opposite Christian Bale. A lighter, more charming side to the actress was displayed in "Marie-Louise, or The Leave" (1995), in which she played a young woman searching for her lover in a crowded train station. Beckinsale delivered a strong turn as the meddlesome orphan taken in by eccentric relatives in the brittle comedy "Cold Comfort Farm" (also 1995). As Flora Poste, she anchored the film and managed to make a busybody character seem charming, and in some ways it was a warm-up for her tackling "Jane Austen's Emma" (BBC/AE, 1996). Although Douglas McGrath's feature version starring Gwyneth Paltrow had opened on American screens first, this version found its partisans who felt it was more faithful to the spirit of Austen.

Capitalizing on the sass and intelligence she had projected in both "Cold Comfort Farm" and "Jane Austen's Emma", Beckinsale shone as an aristocratic med student who falls in with two charming con men (Dan Futterman and Stuart Townsend) in the underrated caper flick "Shooting Fish" (1997). Adopting a flawless American accent, the actress next registered as the bitchy junior publishing executive seeking fun and perhaps Mr. Right in Whit Stillman's "The Last Days of Disco" (1998). The following year, Beckinsale retained the Americanisms to portray a mousy tourist in Thailand who falls for a slick Australian, dragging herself and her traveling companion (Claire Danes) into accusations of drug smuggling in "Brokedown Palace.” After time out for motherhood, she returned to the big screen as Nick Nolte's daughter in the Merchant Ivory adaptation of Henry James' "The Golden Bowl" (2000).

The attractive actress finally had a shot at more mainstream success with two high profile leading roles in 2001. In the big-budget epic "Pearl Harbor,” she was cast as a US Navy nurse who falls in love with a dashing pilot (Ben Affleck) but when news of his death arrives turns to his best friend (Josh Hartnett) for comfort. And Beckinsale was cast opposite John Cusack in the mildly engaging romantic comedy "Serendipity,” playing a woman who believes more in fate than love at first sight and faces a long but seemingly inevitable road to romance. The actress surfaced again in 2003 in the arty indie "Laurel Canyon" as the icy fiancée of an L.A. native (Christian Bale) who returns to his eclectic mother's home in Laurel Canyon, where Beckinsale's character slowly becomes seduced by the sultry Los Angeles lifestyle.

Her highest profile role to date came in "Underworld" (2003), a glossy supernatural thriller with Romeo-and-Juliet overtones, in which Beckinsale played Selene, a vampire embroiled in her kind's long feud with a werewolf clan who falls in love with one of her blood enemies (Scott Speedman). Beckinsale followed up with another action-packed supernatural thriller, teaming with Hugh Jackman for "Van Helsing" (2004), in which she played Anna Valerious, a vampire slayer from a long line committed to ending the reign of Count Dracula who teams with the count's longtime human foe. The actress was better served by her next project, director Martin Scorsese's Howard Hughes glamorous and visually arresting biopic "The Aviator" (2004), in which the actress provided a sultry spark as the fiery film icon Ava Gardner, Hughes' (Leonardo DiCaprio) most challenging, yet sympathetic, paramour.

Beckinsale next revived the vampire Selene for the sequel, “Underworld: Evolution” (2006), as vampires and werewolves battle each other for ultimate control of the undead. As the violence between the two warring factions increases, Selene and her werewolf beau, Michael (Scott Speedman), try to uncover the secrets of the conflict while delving into their own pasts. Despite poor reviews, “Underworld: Evolution” managed to rake in a descent payday. Meanwhile, she starred opposite Adam Sandler in “Click” (2006), a middlebrow comedy about an overworked architect (Sandler) whose life seemingly changes for the better when a strange Bed, Bath and Beyond clerk gives him a universal remote that can pause, rewind or fast-forward anything—barking dog included. But as the remote gets stuck on fast-forward, causing him to miss all the important events in his life, he realizes that it’s probably better to take the bad with the good rather than let his whole life pass before his eyes.

Returning to the horror genre—an apparent favorite for the actress—Beckinsale starred in “Vacancy” (2007), another in a long line of suspense thrillers released in the early part of the century. In this all-too-obvious take on “Psycho,” Beckinsale played the soon-to-be ex-wife of a man (Luke Wilson) forced to spend the night at a seedy motel run by an odd, but seemingly harmless proprietor (Frank Whaley). But the couple soon discovers that the cache of homemade slasher flicks they have found were shot in the very room in which they are staying—both must put aside their differences and work together in order to avoid becoming the next victims of the sadistic filmmakers. While most horror thriller are brushed off by critics as being redundant and tedious, “Vacancy” received its fair share of positive reviews.