Fred and Gunn search for Angel and Cordy who have been missing for three months. The situation is complicated as Connor kills the only eyewitness to Angel's whereabouts. Wesley finally finds Angel and lets him drink his own blood to save him.
Following Wesley's suggestion, Angel consults the "goddess of the lost" who has information about Cordelia's whereabouts. Angel, Fred, and Gunn break into an auction house to retrieve an object only to discover that a sultry cat burglar has her eye on the same property.
Angel, Fred, and Gunn travel to Las Vegas to enlist Lorne's help in locating Cordelia. Once there, they begin to unravel a mystery involving a casino that gambles with futures instead of money.
Cordy inexplicably returns from a higher plane with no knowledge of her past or where she's been.
Fred speaks at a physics symposium about her published article. While addressing the crowd, a portal opens, revealing a demon with a dark secret about her professor.
The group must kill a vampire when a spell intended to return Cordelia's memory lands them back in high school, but Angel becomes a target as the only one to remember that he is, in fact, a vampire.
Angel, Gun, Lorne, and Wesley face up against a beast from the center of the earth bent on creating destruction through earthquakes and fire storms.
Angel discovers that Connor is trapped in Wolfram & Hart when the building experiences a lockdown, while inside, a beast is on a killing spree.
The gang discovers that the one who is able to stop the beast is among them.
The gang decide to get information on The Beast by summoning his former partner and Angel's alter ego, Angelus, whose transformation can only occur during a state of complete happiness.
The gang puts Angelus in a cage while they question him on how to kill The Beast.
The gang tries to retrieve Angel's soul in order to thwart the large entity that is utilizing The Beast.
Angelus feeds on Lilah, and the son, Connor, now vows to kill the father. Wesley decides to get Faith free from confinement in order for her to help him capture Angelus and The Beast.
Angelus continues his quest to find The Beast's master and does battle with Faith, while Cordelia tells Connor to keep their child a secret.
Faith helps capture Angelus when, after feeding on her, he experiences the effects of a lethal drug she injected into herself. With Faith in a coma and Angelus unconscious, Fred brings in Willow to restore Angel's soul.
After learning of Cordelia's pregnancy, Angel and the gang explore the demonic possibilities of the offspring. Gunn helps Gwen crash a black-tie party in order to rescue a girl.
Angel tries to find out from a demon why Cordelia has turned to the dark side and learns that a higher being has orchestrated her pregnancy, his transformation into Angelus, and the birth of Connor, in the efforts to create an ultimate power.
Jasmine, who the now-unconscious Cordelia has given birth to, is mistakenly perceived by the gang as a random woman in trouble. Following her around, they notice she seems to have the power to make everyone worship her, but after being brainwashed themselves, decide she is a threat that must be destroyed.
Fred searches for a way to save Angel and the gang from Jasmine's power.
Angel and the gang escape from Jasmine and hide out underground in the sewers. Meanwhile, Connor learns that Cordelia has disappeared.
Angel finds himself in Jasmine's dimension while Fred, Wesley, and Gunn are being held captive, and must behead a demon and use its powers to stop Jasmine, who plans to declare peace on Earth. Connor finds Cordelia and, despite Angel's best attempts, ends up killing Jasmine and restoring the world's pain.
Angel and the gang are given the chance to run Wolfram & Hart and make use of all its secret information. Meanwhile, Angel finds himself with an opportunity to change a physically and emotionally wayward Connor's future.