Betty finds everything changing as she begins her new job as associate features editor, especially since her boss is her ex-boyfriend, Matt Hartley, who is still unhappy about their breakup.
Betty gets to run her own Mode photo shoot at the United Nations to promote its anti-malaria initiative, "Nothing but Nets," but her big opportunity becomes a baptism by fire when Daniel's neediness takes its toll and Matt openly demoralizes her.
Betty gets a big scoop with Gucci's new designer, Evan York, with the help of Hilda's connected customer, Sammy, but Marc plots to sabotage her, enlisting a clueless Amanda and Matt in his efforts, culminating in a showdown at the restaurant where Ignacio works.
Betty is overwhelmed with her editorial work, so Wilhemina assigns Marc to be Betty's temporary assistant. Betty's story idea to cover the worst jobs in New York goes south when she has to dress up as a hot dog and do a Bollywood dance number.
Matt asks someone else to be his "plus one" at Wilhemina's fund-raiser for Tibetan orphans, and Daniel hires his new BFF Natalie as his assistant, making Betty feel obsolete. Amanda thinks she's Matt's plus one and keeps it a secret from Betty. Daniel discovers that his mother, Claire, has history with Cal Hartley. Meanwhile, Nico secretly plots with her supposedly dead boyfriend to blackmail Wilhemina.
Hilda's bad-boy high school flame and Betty's schoolgirl crush -- Bobby Talercio -- teaches at Justin's school, causing the sisters to both doubt their feelings for "nice guys" -- such as Archie, in Hilda's case.
Daniel lets the leaders from the Community of the Phoenix make decisions at Mode, and Bennett's promises to reunite Daniel with his dead wife prompt Betty to action.
In the steamy Bahamas, the love triangle between Betty, Matt and Amanda explodes, and romantic pairings abound. Things should be dreamy at the Atlantis resort, where even Shakira is staying, but Betty has a nightmarish photo shoot after Willie learns her nemesis, Penelope Graybridge, snagged a coveted job.