Betty's life is in chaos after a series of very unfortunate events, but despite all that's happened, she's still in denial about one thing -- how much Henry's departure has affected her. Meanwhile, Amanda turns to junk food and to Marc in dealing with the revelation that Fey Sommers was her real mother, and Wilhelmina schemes to turn the tragic Meade family events to her advantage.
Betty doesn't know what to do when she finds out Wilhelmina's hunky new bodyguard is taking the job to a whole new level. She's also confused about her own love life when Henry moves back to town and wants to be friends. Meanwhile, Bradford uses Alexis' amnesia to re-write family history, Hilda comes to terms with Santos's death, and Amanda gets a surprising inheritance from Fey Sommers - a dog.
With Henry back in her life, Betty finds herself distracted by yet another guy - Gio, Mode's new sandwich guy, who both annoys and intrigues her. Meanwhile, the Mode-ites prepare for the social event of the season - the Meades' Black and White Ball.
Betty takes steps toward becoming a writer and invests in a class that proves to be so challenging, she tosses her own writing assignment and instead submits a Mode story written by a woman who survived a bear attack.
Much to Henry's distress, Betty tries internet dating and gets asked out on a bowling date. Meanwhile, Marc finds himself attracted to someone who isn't his type, chubby, but charming photographer Cliff. On the home front, Daniel and Alexis steel themselves for a "family" dinner with step-mom-to-be Wilhelmina, and Hilda, lost in her own world, doesn't notice that Justin is ditching school and running with the wrong crowd.
Betty and Henry continue their illicit romance and plan a secret date to "Wicked," but things don't go as planned when Daniel, thinking she's dating Gio, gifts her with tickets for the two of them which Gio refuses to give up. Meanwhile, Daniel woos an important advertiser, a lady with a penchant for younger men.
As Wilhelmina prepares for her big day, all is not quiet on the Mode front when she realizes her fabulous maid of honor, Victoria Beckham, is stealing her thunder.
In the wake of the disastrous Meade/Slater nuptials, Wilhelmina plots her next moves and Mrs. Meade shows up at the Suarez household. Betty tries to make things right with Daniel. Henry uses his charms to get the Mode weekend receptionist to let him in after hours so he can help Betty get the Meade will.
A Wilhelmina-induced crisis at Mode finds everyone scrambling to re-create an issue on deadline. Desperate times call for desperate measures, as Daniel and Betty try to break wild child cover girl out of rehab for a re-shoot, and Alexis tries to charm the printer into extending their time. Meanwhile, Betty struggles with seeing the ghost of Bradford Meade who hopes to goad her into making good on a promise.
Henry reluctantly agrees to go on a double date with Betty and an unlikely couple -- Hilda and Gio. Wilhelmina's attempt to change her image as the queen of mean fails miserably after she tangles with America's most beloved golden girl, Betty White. Meanwhile, Daniel and Alexis compete at paintball to decide who should run the company.
Amanda visits a psychic for insight into her dad's identity. Meanwhile, disturbed about the message Mode is sending to Justin and his classmates after they visit the magazine, Betty pushes Daniel to use "real" models during fashion week. Wilhelmina discovers that she's going to need another woman to carry her baby in their womb.
Betty's new perfume is driving Henry wild and Betty crazy, literally, after she unwittingly begins using a deadly scent concocted by the late Fey Sommers and starts acting very strangely. Meanwhile, Meade matriarch Claire goes on the stand for Fey's murder, a vindictive Amanda withholds important information, and Wilhelmina takes steps to become a member of the Meade clan -- one way or the other.
Betty takes an assignment from Daniel to interview "Phil Roth," thinking it's a sit down with the great novelist, but later learns this Phil Roth is a writer who has penned a series of books about picking up women. Meanwhile, Daniel hooks up with a sexy new lady, Renee, only to learn they have a shocking connection to each other, and Amanda and Marc hatch a plan to reach out to the man they believe is Amanda's biological dad.
Betty's plans to spend a romantic 24th birthday with Henry are sidelined when Charlie drops in unexpectedly. Meanwhile, Wilhelmina does her devious best to spoil Renee's blossoming relationship with Daniel by hinting there's some dark secrets in her sister's past.
An increasingly delusional Renee becomes convinced that Betty is in love with Daniel and therefore is her rival for his affections. Meanwhile, Hilda's new client at the beauty shop turns out to be ex-neighbor and lifetime nemesis Gina Gambarro, who returns to Queens to rub her fabulous life and wealthy doctor husband in Hilda's face.
Betty switches gears with rival Charlie and tries to befriend her by throwing a baby shower. Meanwhile, Hilda confronts Justin's PE teacher when she learns Justin is flunking gym; Daniel unravels after his break-up with Renee, and Christina finally learns who fathered the baby she's carrying for Wilhelmina.
Betty, still missing Henry, fights her attraction to Gio, but later finds herself stuck with him when both volunteer to chaperone Justin's middle school dance. Meanwhile, Hilda tries to get Coach Diaz to notice her; Amanda agrees to do a reality show with her dad, KISS rock legend Gene Simmons; and Daniel feels the heat when Wilhelmina makes her triumphant return to Mode amid a media frenzy.
Just as Gio makes Betty an incredibly romantic offer, Henry pops back into her life with a proposal of his own. Meanwhile, Betty finds herself centerfield at the charity softball game of the year: Mode vs. ELLE Magazine. In other developments, an orphaned boy from France turns up at Mode with an amazing claim, and Hilda learns some shocking news about Coach Diaz.