Betty makes a moving decision about what to do with her life. Meanwhile, Ignacio gets a new job at a fast food joint, and Wilhelmina's manipulations force Daniel to make some changes of his own.
Wilhelmina, prepping her first issue of the magazine as editor, tries to lure Betty over to the dark side by offering her a job at Mode as her assistant. Meanwhile, Daniel discovers he faces a custody battle if he wants to keep Daniel Jr. in the U.S., Hilda's illicit romance with Coach Diaz heats up, and Christina grows suspicious of Stuart's true motives.
An accident leaves the Modies stunned, and Betty tries to piece together what really happened when a police detective comes looking for some answers. Meanwhile, Ignacio learns something shocking about Hilda and the coach.
Gio returns and Betty tries to make amends, but he's not having it. Meanwhile, Daniel struggles with some shocking family news.
Kimmie Keegan asks for Betty's help in finding a job. Daniel tries a matchmaking service with surprising results in his "perfect" match. Justin tries out for a Broadway musical.
Betty finds herself competing with Kimmie who finally shows her true colors. New citizen Ignacio prepares to vote for the first time. Hilda takes advantage of the increased election traffic to promote her shop. Wilhelmina wants to hire a new financial officer whom Daniel can't stand.
Betty works overtime to get the attention of her cute neighbor, struggling musician Jesse. Meanwhile, Betty's life gets even more complicated when a maxed out and suddenly homeless Amanda becomes her temporary roommate. And in other Mode developments, Marc vacillates when Cliff wants them to move in together.
Betty needs Daniel's help to prevent an issue of Mode from shipping that could result in a PR nightmare for the magazine. The problem is, he's unreachable at a company retreat. Meanwhile, Hilda starts a clandestine beauty salon when the city turns down her permit for a legal business, and Connor's fiance meets the Modies.
Competition gets fierce when Betty and Marc both apply to YETI, the Young Editors Training Program that could be their ticket out of assistant-land, but there's only room for one of them. Meanwhile, Justin makes an unlikely friend, Daniel finds himself attracted to Molly and Wilhelmina does her best to get Connor's attention.
Betty is forced to take a page from scam-artist Amanda's book when the two, both cash-strapped, team up for a Mode website article about "How I Blew Ten Grand without Spending a Dime" during a night on the town in NYC they'll never forget.
Betty juggles work and family without much success as she struggles to impress her idol, Jodie Papadakis, the editor running YETI. Things go from bad to worse when she encounters Teri, an assistant at ELLE magazine, just as the long-running Mode/ELLE rivalry heats up.
Betty is torn between work and home in the wake of a family crisis that finds her at odds with Hilda. Meanwhile, "secret" couples Daniel & Molly and Connor & Wilhelmina try to hide their escalating love connections.
Betty contemplates moving home to help her family and save money. But things keep sidetracking her like the fact that sexy neighbor Jesse is suddenly interested in her again. Meanwhile, Daniel tries to hide his tracks as Connor is determined to find out the identity of the mystery man who stole Molly's heart.
Betty tapes a video tribute for Claire Meade's 60th birthday that's very revealing, just not in the way she planned. But the real shocker comes when Betty accidentally leaves the camera on at home and discovers some revealing footage about Ignacio. Meanwhile, Daniel and Wilhelmina have difficulty adjusting to the new significant others in their lives.
Fashion Week brings chaos to the Modies. Betty copes with simultaneously putting on a show for a bizarre new designer named Heinrich and with an assignment from YETI to partner up with Matt, who works at a sports magazine, to learn about each other's jobs. Meanwhile, Wilhelmina fears she's lost her edge in the fashion world because she's been spending too much time with Conner. Daniel tries giving Molly a makeover.
Betty makes a surprising discovery when an assignment from YETI finds her and Marc analyzing Mode's budget for the past year. Meanwhile, she has to deal with increased attention from Matt and the knowledge that old flame Henry has posted some unexpected news on Facebook.
The Suarez family tries to come up with the cash to buy their rental home after their landlord decides to sell. Ignacio enters a TV cook-off with celebrity chef Frankie Burrata. Meanwhile, the Mode staff deals with the shocking aftermath of recent events.
Betty meets Matt's mother but things don't go as she expects. Meanwhile, Wilhelmina gets creative to solve a cash flow problem, Daniel deals with the latest wrinkle in his relationship with Molly, and the Suarez family conspires to set Hilda up with Archie, the promising city councilman from Queens.
Daniel and Wilhelmina hope that an issue of Mode devoted to sex will turn their fortunes around. Meanwhile, when Matt seems a little shy about taking his romance with Betty to the next level, Marc and Amanda team up to give Betty a sexy makeover, and Hilda is uncertain she wants to continue dating Archie.
The Suarez family gets an invitation to meet Matt's famed billionaire dad, Calvin Hartley, at his legendary Easter egg hunt. While Betty sees this as a chance to get closer to Matt, Daniel seizes the opportunity to get Mode a bailout. Meanwhile Wilhelmina suspects that little William might not be her biological son.
Just as it looks like billionaire Calvin Hartley is going to give Mode the bailout it needs so the magazine won't go under, baby William is kidnapped and Betty is dragged into the ensuing drama. Meanwhile Archie asks Hilda to be a part of his latest political campaign.
Betty teams with Matt and Marc for their all-important final YETI project, a photo shoot with Grammy winner Adele, but Matt's slacking off could jeopardize everything they've worked for. Meanwhile, love is everywhere as two wedding proposals are made, but only one couple will walk down the aisle; Claire and Wilhelmina square off over their roles at Mode; and Matt's mom asks Betty for a favor.
When Betty agrees to move in with Matt, Henry pays a visit to NYC, forcing Betty to admit she still has feelings for her former fiancé. Meanwhile, Betty and Marc wait to see if their YETI editor interviews resulted in actual jobs; Daniel's transformation from playboy to devoted husband makes him a media darling; and Wilhelmina cozies up to Victoria Hartley to find out just what kind of connection Claire Meade has with Cal Hartley.
The sudden death of a Mode editor leaves Betty and Marc competing for the same job. Meanwhile, Wilhelmina demands that Claire tender her resignation at a big awards ceremony or she'll reveal Claire's dark secret to Cal Hartley; Matt lets his jealousy over Henry get the best of him; and Justin nervously waits to see if he got into performing arts school.